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(1)

783

Copyright © 2015 The Korean Society of Fisheries and Aquatic Science pISSN:0374-8111, eISSN:2287-8815

서 론

진도주변연안은외해역에서유입되는고온

,

고염의대마난 류와계절적으로유입되는저염의양자강담수

,

서해연안수 다양한해류와연안육수의영향으로복잡한해양환경을나타 내고있다

(Jung et al., 2012).

그리고강한조류가작용하는 역적인특징때문에진도연안역은낭장망

,

안강망

,

주목망

수동적인어구를이용한어업이발달해왔다

(NFRDI, 2005).

낭장망어업은조류의흐름을이용하여유영능력이약한어류

,

갑각류등을어획하며

,

경제적가치가높은멸치

(Engralius ja- ponicus)

주요대상종이다

(Cho et al., 2014).

남해주변연안에서낭장망을이용한연구들을살펴보면

,

부분주요어업대상종인멸치가전체어획량의

90%

이상으로 높은출현량을 보인다

(Han and Oh, 2007; Shin et al., 2013;

Yoo et al., 2014).

하지만낭장망은망자루끝에세목망을설치

하여멸치를제외한기타소형어와치어

(Juvenile)

어획사망

율이높아기아

(inanition),

포식압

(predation pressure),

환경요

(environmental factor)

자연사망과함께초기사망에영향

미치는하나의요인으로여겨지고있다

.

그래서낭장망에 집된어류종조성을분석한다면

,

낭장망어업에관한적절한 책과규제를설정하는데도움이있을것으로생각되었다

.

연구의목적은진도주변연안에서낭장망에의해채집된 어류종조성파악과함께주요대상종인멸치를비롯한부수어 획종

(

소형어류

,

치어

)

월변동을확인하고

,

채집된주요어류 주거종

(redident species)

회유종

(migratory species)

으로 구분하여향후

,

지속가능하고효율적인어업관리방안을 시하고자한다

.

재료 및 방법

연구에 사용된 시료는 전남 진도 연안

(34

°

22’51.30”N, 126

°

6’50.16”E)

에서

2014

5

월부터

11

월까지매월

1

전체 길이

64 m,

끝자루망목크기

4.0 mm

낭장망

(gape net with

wings)

어획된어획물을매월

5 kg

무작위로표본채집하

였으며

,

시료의부패를방지하기위하여쿨러

(cooler)

빙장

보관하여연구실로운반하였다

(Fig. 1).

채집된어류의동정은

진도 연안 낭장망에 어획된 어류의 종조성

정재묵·유준택·김희용 1 ·이선길·고우진·김영혜*

국립수산과학원 남해수산연구소 자원환경과, 1국립수산과학원 연구기획과

Species Composition of Fish Collected by a Gape Net with Wings in the Coastal Waters of Jindo, Korea

Jae Mook Jeong, Joon-Taek Yoo, Heeyong Kim

1

, Sun-Kil Lee, Woo-Jin Go and YeongHye Kim*

Fisheries Resources and Environment Division, South Sea Fisheries Research Institute, National Institute of Fisheries Science, Institute of Fisheries Science, Yeosu 59780, Korea

1

Research and Development Planning Division, National Institute of Fisheries Science, Busan 46083, Korea

Species composition of the fish in the coastal waters of Jindo was determined using monthly sample collection by a gape net with wings in 2014. Of a total of 41 fishes collected, the dominant species were Engraulis japonicus , Thryssa kammalensis , Sardinella zunasi , Leiognathus nuchalis , Amblychaeturichthys hexanema , Neosalanx ander- ssoni , Acanthogobius flavimanus . The former three species were the migratory species and the other four species were the resident species. These 7 fishes accounted for 99.6% of the total number of individuals collected. Monthly species composition did not show a clear seasonal trend. The peak number of individuals occurred in May, lowest in November. Fish were divided into four groups by the cluster analysis.

Key words: Jindo, Gape net with wings, Species Composition, MDS

This is an Open Access article distributed under the terms of the Creative Commons Attribution Non-Commercial Licens (http://creativecommons.org/licenses/by-nc/3.0/) which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original work is properly cited.

http://dx.doi.org/10.5657/KFAS.2015.0783 Korean J Fish Aquat Sci 48(5) 783-788, October 2015

Received 18 August 2015; Revised 18 September 2015; Accepted 29 October 2015

*Corresponding author: Tel: +82. 61. 690. 8941 Fax: +82. 61. 686. 1588

E-mail address: [email protected]

(2)

Chyung (1977), Nakabo (2002), Kim et al. (2005)

이용하였 으며

,

종별로개체수를계수하였다

.

조사해역의수온을측정하

위하여매월어구에소형수온측정계

(HOBO Water Temp

Pro v2)

이용하였다

.

통계분석을위한자료의정규화

(normality)

분산을동질화

(homocedasticity)

시키고우점종의

bias

줄이기위하여어류 출현량의 로그변환

[logarithmic transformations, log

10

(x+1)]

수행하였다

.

월별어류군집의변화를분석하기위하여분산 분석

(one-way ANOVA)

실시하였고

, Tukey test

검정하였

.

50

개체이상출현한

7

종의출현시기에대한유사도를확인하 위하여

Pianka (1973)

중복도지수를이용하였다

.

A

ij

=

∑(Pih - Pjh)

∑P2ih×∑P2jh

(P

ih

:

채집시기

h

채집된전체개체수에대한어종

i

개체 수의비율

P

jh

:

채집시기

h

채집된전체개체수에대한어종

j

개체 비율

)

구해진유사도를이용하여출현시기의유사성을나타내기 하여

dendrogram

다차원척도법

(MDS, Multi-dimensional scaling)

이용하였다

.

통계분석을위하여

PRIMER v5 statis- tical package

이용하였으며

(Clarke and Gorley, 2001),

이때 비우점종

(50

개체이하

)

오류를증가시킬확률이높기때문에 분석에서제외시켰다

.

결 과

수온

조사해역의수온은

16.6-20.9℃

범위를보였으며

,

봄철인

5

월에가장낮았고여름철인

8

월에가장높았다

.

출현종

조사기간동안

13

29

41

67,845

개체의어류가

집되었다

(Table 1.).

분류군별출현어종수를살펴보면농어목

(Perciformes)

12

15

종으로 가장 많이 출현하였으며

,

다음으로 청어목

(Clupeiformes) 2

8

,

쏨뱅이목

(Scorpae- niformes) 3

3

,

바다빙어목

(Osmeriformes) 1

3

,

귀목

(lophiiformes),

가자미목

(Pleuronectiformes),

뱀장어목

(Anguiliformes)

각각

1

2

종이출현하였다

.

복어목

(Tetraodoniformes),

홍매치목

(Aulopiformes),

대구목

(Gadi- formes),

샛비늘치목

(Myctophiformes),

큰가시고기목

(Gas- terosteiformes),

숭어목

(Mugiliformes),

바다빙어목

(Osmeri- formes)

1

1

종씩출현하였다

.

개체수에서가장많이출현한어종은멸치

(Engraulis japoni- cus)

64,700

개체채집되어전체출현개체수의

95.4%

차지하였다

.

다음으로는청멸

(Thryssa kammalensis)

댕이

(Sardinella zunasi)

이가 각각

1,372, 1,096

개체가채집되 전체개체수의

2.0, 1.6%

차지하였다

.

주둥치

(Leiog- nathus nuchalis),

도화망둑

(Amblychaeturichthys hexanema),

도화뱅어

(Neosalanx anderssoni),

문절망둑

(Acanthogobius flavimanus)

순으로채집되었고

,

전체채집개체수의

99.6%

차지하여 우점하였다

.

볼락

(Sebastes inermis),

전갱이

(Trachurus japonicus),

붕장어유생

(Conger myriaster lepto- cephalus)

등이출현하였다

.

월별 출현종

월별종수는

5

월에

22

종으로가장많았고

, 11

월에

7

종으로 적었다

(Table 1.).

10

월에는

9

종이채집되어비교적 었고

, 6-9

월에는

12-14

종이채집되어비슷한종수를보였다

.

출현개체수는

7

35,600

개체로가장많았으며

, 8

1,369

개체로가장적은개체수를 보였다

. 5

, 6

, 10

월에는각각

6,055, 6,024, 6,380

개체로비슷했으며

, 9

월과

11

월에는각각

7,767, 4,648

개체가채집되어월별출현개체수는유의한차이

보였다

(ANOVA, P<0.05).

최우점종인멸치는채집기간동안지속적으로높은출현량을 보였으며

,

특히

7

월에

35,454

개체로극우점하였다

.

아우점종 청멸은

5-8

월까지는

9-16

개체로적은출현량을보이다가

, 9

월부터

11

월까지

147-925

개체로높은출현량을보였다

.

밴댕 이와주둥치는

10

월과

9

월에각각

820, 122

개체로높은출현량 보였다

.

도화망둑과문절망둑은각각

7

월과

6

월에각각

88

, 58

개체로집중적인출현양상을보였으며

,

도화뱅어는

8

60

개체로높은출현량을보였다

(Table 1.).

Fig. 1. Sampling station (a black circle) for the study in the coastal waters of Jindo, Korea.

123124125126127128129130131132 31 32

33 34 35 36 37 38

125° E 125.5° E 126° E 126.5° E 127° E

34° N 34.5° N 35.5° N

35° N 36° N

Jindo Island

Jeollanam -do

S.

0.8 1.0 0.6

0.4 0.2

0

Similarity

GroupⅠ

GroupⅡ

GroupⅣ GroupⅢ

L. nuchalis A. flavimanus A. hexanema N. anderssoni E. japonicus

zunasi T. kammalensis

Sn Ty

Ej Na

Af

Ah Ln

Stress : 0.01

GroupⅠ GroupⅡ

GroupⅢ

GroupⅣ

(3)

Table 1. Species composition of fish collected in the coastal waters of Jindo, Korea (+: less than 0.1) Month

Scientific name

May Jun. Jul. Aug. Sep. Oct. Nov. Total

N N N N N N N N %N

Engraulis japonicus 5,965 5,911 35,454 1,283 7,033 5,376 3,678 64,700 95.4

Thryssa kammalensis 16 3 11 9 261 147 925 1,372 2.0

Sardinella zunasi 1 6 252 820 17 1,096 1.6

Leiognathus nuchalis 1 1 122 32 21 177 0.3

Amblychaeturichthys hexanema 88 1 1 90 0.1

Neosalanx anderssoni 1 60 3 64 0.1

Acanthogobius flavimanus 2 58 60 0.1

Sebastes inermis 29 7 36 0.1

Trachurus japonicus 13 11 4 1 29 +

Conger myriaster leptocephalus 3 1 1 22 27 +

Mugil cephalus 25 25 +

Siganus fuscescens 22 22 +

Saurida elongata 22 22 +

Salangichthys microdon 16 16 +

Upeneus japonicus 2 5 8 15 +

Sphyraena pinguis 1 14 15 +

Trichiurus lepturus 2 6 1 2 1 12 +

Foetorepus altivelis 11 11 +

Thryssa adelae 1 1 4 1 7 +

Pampus echinogaster 6 6 +

Konosirus punctatus 3 3 6 +

Pseudaesopia japonica 4 4 +

Conger myriaster 4 4 +

Clupea pallasii 4 4 +

Collichthys niveatus 1 2 3 +

Syngnathus schlegeli 1 1 2 +

Setipinna tenuifilis 2 2 +

Salanx ariakensis 1 1 2 +

Bregmaceros japonicus 1 1 2 +

Benthosema pterotum 1 1 2 +

Tridentiger barbatus 1 1 +

Takifugu pardalis 1 1 +

Ophichthus urolophus 1 1 +

Lophius litulon 1 1 +

Histrio histrio 1 1 +

Cynoglossus joyneri 1 1 +

Cryptocentrus filifer 1 1 +

Cottidae 1 1 +

Coilia nasus 1 1 +

Chelidonichthys spinosus 1 1 +

Champsodon snyderi 1 1 +

Chaetodon modestus 1 1 +

Total 6,055 6,024 35,600 1,369 7,769 6,380 4,648 67,845 100.0

(4)

출현종에 대한 출현시기 유사성

출현종에대한출현시기유사성비교는

7

종을대상으로실시 하였으며

,

중복도지수를구한

, cluster

분석을이용해

dendro- gram

다차원척도법

(MDS)

으로분석한결과유사도

80%

준에서

4

그룹으로구분되었다

(Fig. 2). GroupⅠ

여름과 을에출현량이높았던그룹으로멸치

,

청멸

,

밴댕이가속하였으

, Group Ⅱ

여름에출현량이높았던그룹으로도화뱅어가

속하였다

. Group Ⅲ

여름철

7

월과

6

월에출현량이높았던 그룹으로도화망둑과문절망둑이속하였으며

, Group Ⅳ

가을에출현량이높았던그룹으로주둥치가속하였다

. 소형어 및 치어의 출현

낭장망에채집된어류

, 10

개체이상출현한소형어와치어

Table 2.

같다

.

소형어는청멸

,

밴댕이

,

도화망둑등이치어 로는갈치

,

볼락

,

붕장어

,

숭어

(Mugil cephalus),

전갱이

,

문절망 등이출현하였다

(Table 2.).

고 찰

조사가이루어진진도주변해역의수온은여름인

8

월에도

21℃

이하로일반적인남해연안과비교하였을

,

비교적낮았

.

이는여름철진도주변해역에황해저층냉수영향을받은 강한냉수대가우리나라남서해역에조석전선을형성하였기 때문인것으로판단되었다

(Jeong et al., 2009).

기존에낭장망을이용한선행연구를살펴보면

,

남해도주변 해역에서는

56

(Huh and kwak, 1998b),

서해고군산군도에 서는

53

(Hwang, 1998),

여수연안에서는

63

(Han and Oh, 2007),

진도수품리연안에서는

53

(Shin et al., 2013)

채집 되어연구

(41

)

비교하였을

,

비교적많은종이채집된 것을있었다

.

이와같이종수에서차이를나타낸이유는

,

수동적어구인낭장망의어획특성과연구마다다른어구의 크기

(

남해도

: 10 m;

고군산군도

: 15 m;

여수

: 100 m;

진도

: 70

m,

가로길이

),

해역마다가지는수괴의특색다양한요인에

의한것으로판단된다

.

조사기간동안멸치

(E. japonicus)

64,700

개체로조사가 시작된

5

월부터종료시점인

11

월까지지속적으로우점출현하 였는데

,

이는낭장망어업의주요목표종이멸치이고

,

군집을 루는특성과연중지속적인산란을통해많은개체수가우리나 연안에분포하기때문인것으로판단되었다

(NFRDI, 2005).

하지만진도에서과거수행된연구

(Shin et al., 2013)

비교해 보면

(Table 3.),

멸치의어획량이확연히감소하였는데

,

이는 분석방법과무작위로채집된시료의양에서오는차이인것으 추측되지만

,

다른환경요인과어란

·

치어의분포등도 고려해야것으로생각된다

.

청멸

(T. kammalensis)

밴댕이

(S. zunasi)

각가

1,372, 1,096

개체가채집되어우점하는양상 보였는데

,

이는연안에서청멸과밴댕이가무리를이루어

식하는생태적특징과유영수층이

·

중층인부어류

(pelagic

fish)

이기때문인것으로추측되었다

(Kim et al., 2005).

청멸은낭장망을이용한연구이외에도남해안주변 안에서다른어구를이용한연구에서도우점하는것으로보고 되었다

(Huh and Kwak, 1998a; Lee et al., 2011; Jeong et al., 2013).

다음으로개체수가많은주둥치

,

도화망둑

,

문절망둑 진도주변연안에서식하는주거종

(resident species)

으로 Table 2. Growth stages of fish caught by a gape net with wings in the coastal water of Jindo, Korea

Small fish Juvenile stage Sardinella zunasi Trichiurus lepturus Leiognathus nuchalis Sebastes inermis

Thryssa kammalensis Conger myriaster leptocephalus Amblychaeturichthys

hexanema

Mugil cephalus Trachurus japonicas Acanthogobius flavimanus

Fig. 2. A dendrogram and multi-dimensional scaling (MDS) analy- sis of 7 most numerous species collected by a gape net with wing in the coastal water of Jindo, Korea. (Ej: Engraulis japonicus, Ty:

Thryssa kammalensis, Sn: Sardinella zunasi, Ln: Leiognathus nu- chalis , Na: Neosalanx anderssoni, Ah: Amblychaeturichthys hex- anema, Af: Acanthogobius flavimanus)

123124125126127128129130131132 31 32

33 34 35 36 37 38

125° E 125.5° E 126° E 126.5° E 127° E

34° N 34.5° N 35.5° N

35° N

Jindo Island

Jeollanam -do

S.

1.0 0.6 0.8

0.2 0.4 0

Similarity

GroupⅠ

GroupⅡ

GroupⅣ GroupⅢ

L. nuchalis A. flavimanus A. hexanema N. anderssoni E. japonicus

zunasi T. kammalensis

Sn Ty

Ej Na

Af

Ah Ln

Stress : 0.01

GroupⅠ GroupⅡ

GroupⅢ

GroupⅣ

(5)

측되었는데

,

이는주로저층가까이서식하거나저질에기대어 서식하는반저서어류

(semi benthic fishes)

저서어류

(benthic fishes)

이기때문인것으로판단되었다

(Nakabo, 2002; Kim et al., 2005).

외에적은개체수가채집되었지만

,

볼락

(S. inermis),

전갱

(T. japonicus),

붕장어유생

(C. myriaster leptocephalus),

(M. cephalus),

독가시치

(Siganus fuscescens)

등은모두 어가아닌자어

(larvae)

또는치어

(juvenile)

였는데

,

볼락과독가 시치는생태적특성상주거종으로진도주변연안에서부화한 것으로판단되지만

,

전갱이

,

붕장어유생

,

숭어등은회유종

(mi- gratory species)

으로산란장과부화장소의추측이어려우며 해류에의해조사해역으로수송되어채집된것으로생각 되었다

.

채집어종수의월변동을살펴보면

, 5

월에가장많았으

, 11

월에가장적은종수를보였는데

,

이는수온이가장 향을미친것으로판단된다

(Kim et al., 2002).

최우점어류인 치는채집기간동안지속적으로많은개체수를보였는데

,

특히

7

월에가장많은

35,454

개체가채집되었다

.

이는멸치의주요 란시기인초여름에부화한멸치가진도주변연안해역으로 가입

(recuritment)

되었기때문인것으로판단되었다

.

청멸과 밴댕이는

9

월부터멸치의개체수가줄어들면서증가하는양상 보였는데

,

이는연안생태계내에서같은부어류로구분되는 어류사이의

·

공간분포

(temporal and spatial distribution)

작용한하나의생태적기작인것으로판단된다

(Wotton, 1990).

cluster

분석을실시한결과

, 4

개의그룹으로나눌있었는데 멸치

,

청멸

,

밴댕이는집중적으로출현한시기가유사하였으며

,

도화망둑과문절망둑은일정하지못한출현량을보였다

.

문절 망둑의산란기는

2-3

월로알려져있는데이와같은연구결과

,

늦겨울에서초봄에부화한개체들이치어로성장해

6

월에집중적으로채집된것으로추측된다

(Park et al., 2005).

문절망둑은채집되지않았는데

,

이는성장에따른형태 발달이수반됨에따라낭장망에이상채집되지않은 으로추측된다

.

앞서언급한바와같이낭장망어업은멸치를어획대상으 하지만

,

소형어류와다양한어류의

·

치어의혼획이발생하 적절한관리방안과대책의필요성이항상언급되어왔다

.

,

지속적인낭장망어업을위해서는장기간모니터링을통한 금어기설정

,

어업규제강화와같은정책체계가확립되어야 것으로생각된다

.

사 사

논문은

2015

년도국립수산과학원수산과학연구사업

(

낭장

어업의합리적관리방안연구

, RP-15-FE-28)

지원으로 행된연구이며

,

연구비지원에감사드립니다

.

References

Cho SG, Song DH, Yoo JT, Choi MS and Cha BJ. 2014. Com- parison of fishing gears and catches by areas in the gape net with wings in Jeonnam, Korea . Bull Korean Soc Fish Tech 50, 301-309. http://dx.doi.org/10.3796/KSFT. 2014. 50. 3.

Chyung Mk. 1977. The fishes of Korea. Ilji-sa, Seoul, Korea.301.

Clarke KR and Gorley RN. 2001. Getting started with PRIMER v5: User Manual/Tutorial. Primer-E, Plymouth.

Han GH and Oh YS. 2007. Species composition and quantitative fluctuation of fishes collected by gape net in coastal waters Table 3. Comparison of major species between previous and present studies in the coastal waters of Jindo, Korea

Source Present study Shin et al. (2013)

Study period 2014 2011

Sampling gear gape net with wings gape net with wings

Species N (ind.) Rank N (ind.) Rank

Engraulis japonicus 64,700 1 546,219 1

Thryssa kammalensis 1,372 2 16,952 2

Sardinella zunasi 1,096 3 5,147 5

Leiognathus nuchalis 177 4 235 14

Amblychaeturichthys hexanema 90 5 - -

Neosalanx anderssoni 64 6 - -

Acanthogobius flavimanus 60 7 - -

Ammodytes personatus - - 8,514 3

Muraenesox cinereus - - 6,180 4

Collichthys lucidus - - 4,603 6

Konosirus punctatus 6 24 3,125 7

(6)

of Yeosu, Korea. Bull Korean Soc Fish Tech 43, 261-273.

Hwang SD. 1998. Diel and Seasonal variations in species com- position of fishery resources collected by a bag net off Kog- unsan-gundo. Ko rean J Ichthyol 10, 155-163.

Huh SH and Kwak SN. 1998a. Seasonal variation in species composition of fishes collected by an otter trawl in the coast- al waters off Namhae Island. Ko rean J Ichthyol 10, 11-23.

Huh SH and Kwak SN. 1998b. Seasonal variation in species composition of fishes collected by winged stow nets on an- chors off Namhae Island. Bull Korean Soc Fish Tech. 34, 309-319.

Jeong HD, kwoun CH, Kim SW and Cho KD. 2009. Fluctua- tion of tidal front and expansion of cold water region in the Southwestern sea of Korea. Korean J Mar Environ Safety 15, 289-296.

Jeong JM, Park JM, Huh SH, Ye SJ, Kim HJ and Baek GW.

2013. Seasonal variation in the species composition of fish assemblages in the coastal waters off Gadeok-do, South Sea, Korea. J Korean Fish Soc 46, 948-956. http://dx.doi.

org/10.5657/KFAS.2013.0948.

Jung SW, Park JG, Jung DH and Lim DI. 2012. Seasonal Changes in Water Masses and Phytoplankton Communities in the Western Part of South Coastal Waters, Korea. Korean J Environ Biol 30, 328-338. http://dx.doi.org/10.11626/

KJEB. 2012. 30. 4. 328.

Kim JK, Choi OI, Chang DS, Kim JI. 2002. Fluctuation of Bag- net Catches off Wando, Korea and the Effect of Sea Water Temperature. J Korean Fish Soc 35, 497-503.

Kim IS, Choi Y, Lee CH, Lee YJ, Kim BJ and Kim JH. 2005.

Illustrated Book of Korean Fishes. Kyohak Publ., Seoul, Korea.

Lee SG, Seo YG, Kim JI, Kim HY and Choi MS. 2011. Sea- sonal Species Composition and Fishes By Beam Trawl in Yeoja Bay. Ko rean J Ichthyol 23, 206-216.

Nakabo T. 2002. Fishes of Japan with Pictorial Keys to the Spe- cies. Tokai Univ. Press.

NFRDI (National Fisheries Research and Development Insti- tute). 2005. Commercial Fishes of the Coastal and Off shore Water in Korea. Hangul Graphics Busan, Korea.

Park MH, Hwang IJ, Kim DJ, Lee YD, Kim HB and Baek HJ.

2005. Gonadal Development and Sex Steriod Hormone Levels of the Wellow fin Goby Acanthogobius flavimanus. J Korean Fish Soc 38, 309-315.

Pianka ER. 1973. The structure of lizard communities. Ann Rev Ecol Syst 4, 53-74.

Shin JG, Choi MS, Seo Yl, Cha HK, Lee SG, Kim HY and Oh TY. 2013. Study for Fishery Management measure on Gape Net with Wings with Catch Composition in the Water of Jin- do. Bull Korean Soc Fish Tech 49, 404-418. http://dx.doi.

org/10.3796/KSFT. 2013. 49. 4. 404.

Wotton, R.J. 1990. Ecology of Teleost Fishes. Champman Hall, New York, USA

Yoo JT, Kim JG and Choi MS. 2014. Change of Structure Com- munity of Fish Collected by a Gape Net with Wings after 12 Years in the Coast of Wando Island, Korea. J Korean Fish Soc 47, 659-666. http://dx.doi.org/10.5657/KFAS.2014.0659.

수치

Fig. 1. Sampling station (a black circle) for the study in the coastal  waters of Jindo, Korea.
Table 1. Species composition of fish collected in the coastal waters of Jindo, Korea (+: less than 0.1) Month
Fig. 2. A dendrogram and multi-dimensional scaling (MDS) analy- analy-sis of 7 most numerous species collected by a gape net with wing  in the coastal water of Jindo, Korea

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