Distribution of Meiobenthic Communities in the Deep-sen Floor of Northeastern Pacific Seafloor along a Latitudinal Transect

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(1)Ocean and Polar Research. Vol. 26(2):255-263. June 2004. Article. §ÿ ï·

(2) ö *~º 7;*ÿ> `÷~ *êê ª

(3) ß9. ö8 ÁBÿ9 ÁBë* 1*. 1. 2. ]·\ö ·>¶ö\¦ ]·\ö ~ãÁ¶ö\¦ (425-600) ã8ê nÖ nÖÖÚ] Ò* 29^ 1. 2. Distribution of Meiobenthic Communities in the Deep-sea Floor of Northeastern Pacific Seafloor Along a Latitudinal Transect Won-Gi Min1* , Dong-Sung Kim1, and Woong-Seo Kim2 1. Marine Living Resources Research Division, KORDI Marine Geoenvironment and Resources Research Division, KORDI Ansan P.O. Box 29, Seoul 425-600, Korea. 2. Abstract : This study was conducted to investigate the community structure and distributional pattern of meiobenthos in the deep-sea bottom of the Clarion-Clipperton Fracture Zone of the northeastern Pacific during July 2003. The samples were collected using the multiple corer at 13 stations along the transects from 5oN to 17oN. The most abundant meiobenthos was nematodes and followed by sarcomastigophorans; these two taxa comprised more than 70% of the total abundance at all stations. For size distribution analyses showed that animals which fit into the sieve mesh size of 0.063 mm were abundant. The maximum density of meiobenthos was 147 inds./10 cm2 at station located in 7oN, and the minimum density was 6 inds./ 10 cm2 at station located in 14oN. More than 60% of meiobenthos were distributed at surface sediment layer within 1.0 cm, and the peak abundance was found at 0~0.25 cm layer. The latitudinal distribution pattern of meiobenthos in the study area seemed to be related with the primary productivity of the surface water that is also connected to the water circulation pattern of the Pacific Ocean near the Equator, diverging at latitude of 8 oN and conversing at 5oN..

(4) . 7;*ÿ>. Key words : (Meiofauna), Pacific), (Deep-sea bed). 1. *. `÷\`(Community. . º *Ú ·öB Ö 'f 'j Næ~ ®æ ò 7~8& ÚJÚ F jL' Ì² \& Ú ^, 20^8 ê>ö B¢ öÒ8. Ëj~ BB ö B~º >ö Ï' 7 \& ·>î, ß® Multiple Aberdeen corerf ?f BïB Ëj~ BB îÚ þO»~ 'Ïf ö *Corresponding author. E-mail : [email protected]. structure),. §ÿï·(Northeastern. & 'j æzÊº êV¢ îN "Vê ~&. ~ '>ö B~º 7;&Bÿ>f &;&B ÿ>ö j V& Ö ·, * Ú7 ÒW ¸b, Öïê V r^ö >~ Ã&ö V b ï 6² VÞV& ² jò Þ (Gerlach 1971; Shirayama 1994). r^ö &BêöB 7;& Bÿb~ 7ºWf ç&'b nö j~ ² ¦' >Ú, f &NB & " Ú B~² Úæ ® . 1970j *ö >¯B &» ~~ 7;& (Giere 1993)..

(5) Min, W.-G. et al.. 256. Bÿbö & º 1964jö §jzÒ~ n~ & Ò ç¦öBf 1966j ÿj*Ò~ &öB~ & *¦& (Soltwedel 2000). ê · ~ã ~ 9f æöB ' &·j &çb 7;&Bÿb ö & ¶ò& »'>V ·>îº, " &B· " æ7ö & & "¢ îb(Gooday and Lambshead 1989), ï·~ º " Bï· öB 7;&Bÿb~ ªßW" ~ãºö & Þ ~ (Shirayama 1983, 1984a, 1984b, 1990; Snider et al. 1984; Alongi and Pichon 1988; Alongi 1992)f, §ÿ ï·öBº " Ú &7b¶ö jf &NB &[ ê~ B~ãæzf v¦ö & 'Ëj ï&~º & Úr (Renaud-Mornant and Gourboult 1990; Thiel and Schriever 1993; Ahnert and Schriever 2001; Lambshead et al. 2001, 2003). , (2002) , , 1994 UN CCFZ(Clarion-Clipperton Fracture Zone) ( 1997; 2000; 1996, 1997, 1998, 1999, 2000, 2001, 2002, 2004). CCFZ , ,. Þ Ú öB Úê 7;&Bÿbö & º f >¯ ëê ¦" &ÒöB ~ ª Þj B~º *Z ; æ ö &~º j b¦V j7j * A f §ÿ ï·~ öB 7;&Bÿb~ ª & >¯>Ú z f f ·>Ö¦ ~ &çæ º 'ê¢ 7b v º Î'ê ~¾ §'ê >~ §'ê ~& ê.ö V ¢ $º Vö V¢ º*& Fÿ'b æ~º æ b, æz ·çö V¢ >7~ b¾ ö 'Ëj Aº &Bb~ ª 5 B·çê ¢î > ® . æ, "æ ö & 7º* Òº Nê *Z j7æöB~ j7 ±f ¶ò, "æ ~ ãæz~ ªC, ¯ ·~ *Ú' FV' ç&&ê ¢ 2k~º ö êæ >Ò¢ 'B . *Òræ CCFZöB >¯B 7;&Bÿb~ º " j7æ CCFZ~ §* 6~16 º* ÚöB ¦ª'b >¯> îº, º ¦ z 7º* æ~ 7;&Bÿb ~ ÷ ßWj *êê 2k~¶ §* 5~17 ~ Ò æFj 7b ' *êê ª ßWj ªC~¶ ~ & . o. o. 2. Òò 5 O». & 7 æ~ §* 5êöB 17êö º Ò æFj 7b 7;&Bÿb~ B&ê, bç, Ú ï, >ç' ªßW ~ ' ßWj ªC~V *~, §* 5êöB 17êö Vræ ' *ê ;6 N05, N06, N07, N08, N09, N10, N11, N12, N13,. Fig. 1. A map showing the study area and sampling stations for meiobenthos near the CCFZ of Northeastern Pacific during July 2003.. ~ C 13B Ò;6öB 7;&B bò¢ j÷~& (Fig. 1). òjº 2003j 7ú 3 ¢¦V 8ú 1¢ræ ·ö~ ·«ÒF N *Ò^¢ Ï~ M.C.(Multiple corer) &~ 'b j j~&b, 7 ;6 N11, N15öBº [ö ª ~º ï~ *Z~ 'Ëb ;ïò jö N ~&, ;;òò j÷ &Ë~&, ¾^æ 11B~ ;6 öBº ;ï 5 ;Wò¢ j~& (Table 1). * ;6 öB j÷B 'b¦V 7;&Bÿbj j~V *~ ã 3.6 cm V~ jÚ zÚ¢ ÒÏ~ ¦ò¢ j~& . æÂ j*~ Ö" (f 1997; f 2000)b¦V 7;&Bÿb 'b~ [öBº j". · æz¢ ®Ú ¢ 76'b Ò~ V *~, ò¢ [f ^ªz~ C 15B [ ¯, 'b~ [b¦V 1 cmº 0.25 cm vþ . ~(0~0.25, 0.25~0.5, 0.5~0.75, 0.75~1.0 cm), 1~3 cm N14, N15, N16, N17.

(6) Meiobenthic Community of Deep-sea Bed in the Northeastern Pacific. 257. Table 1. The information of the sampling station in Northeastern Pacific. Station. Location (N) (W). Depth. N05. 05o00.937 131o30.000 4324 m. N06. 06o01.940 131o30.079 4090 m. N07. 07o01.790 131o30.952 4601 m. N08. 08o01.771 131o30.011 5080 m. N09. 08o58.959 131o31.292 4871 m. N10. 09o59.539 131o30.100 4945 m. N11. 11o01.301 131o29.226 4938m. N12. 12o00.018 131o30.236 5015 m. N13. 13o01.268 131o29.206 4964 m. N14. 14o00.124 131o30.199 4997 m. N15. 15o00.120 131o30.108 5006 m. N16. 15o58.147 131o31.064 5035 m. N17. 16o58.784 131o31.025 4833 m. Time (G.M.T.) 2003/07/25 10:54 2003/07/24 16:39 2003/07/23 06:37 2003/07/22 14:28 2003/07/21 06:15 2003/07/19 14:35 2003/07/09 22:16 2003/07/11 09:24 2003/07/12 06:21 2003/07/13 09:53 2003/07/14 13:16 2003/07/15 03:43 2003/07/16 14:42. Gear. Label. Sample Volume. Fixative. MC Quan. A/B~B/B 10 cm2 ×1 cm×10 5% rose bengal formalin sol. MC MC MC MC MC. Quan. A/B~C/C Qual. A/A Quan. A/B~C/C Qual. A/A Quan. A/B~C/C Qual. A/C~B/B Quan. A/B~B/B Qual. A/C~B/B Quan. A/B~B/B Qual. A/C~B/B. 10 cm2 ×1 cm×10 50 cm2 × 5 cm 10 cm2 ×1 cm×10 50 cm2 ×5 cm 10 cm2 ×1 cm×10 10 cm2 ×10 cm 10 cm2 ×1 cm×10 10 cm2 ×10 cm 10 cm2 ×1 cm×10 10 cm2 ×10 cm. MC Qual. A/C~C/C 50 cm2 ×5cm Quan. A/C~C/C Qual. A/C~C/C Quan. A/C~C/C MC Qual. A/C~C/C Quan. A/A MC Qual. A/C~C/C MC. 10 cm2 ×1 cm×15 10 cm2 ×10 cm 10 cm2 ×1 cm×15 10 cm2 ×10 cm 10 cm2 ×1 cm×10 10 cm2 ×10 cm. MC Qual. A/D~D/D 10 cm2 ×10 cm Quan. A/B~B/B Qual. A/C~C/C Quan. A/B~B/B MC Qual. A/C~B/B MC. ræ~ 'bf 0.5 cm vþ ªÒ~&b(1.0~1.5, 1.5~2.0, 2.0~2.5, 2.5~3.0 cm), ¾^æ òº 1 cm vþ ¾¢B ªê j~& . ''~ [b . 'b ò f ®Û.(Rose Bengal)Wî "ïB¢ b 5% 7W öÖb ;® . Ú>>Úê b òº öB ' V~ Ú(1 mm, 500 µm, 250 µm, 125 µm, 63 µm, 37 µm) Vê ÚÚ 7 *ã ~öB ª~ 5 ê>~& . 'bö B~º 7;&Bÿb~ &ê 5 >çªº 10 cmræ ¾Ò ò~ ªCöB * b~ Â*BÚ>& [ 3 cmn ö 90% çj Næ~ ®V r^ö öBº 3 cmræ~ ªC ¶ò¢ ÒÏ~ ' ;6~ & 8b *~& . $ *ã Òê" 'j *~ &Ò ^Ö, öêR, Ã~>¢ b Ïb ;j ~~ B Vö IÚ BB® ÃBÎ êö, BÛV nö I Ú j*® >ªj ÃBBB, H-S Ò¢(Shirayama et al. 1993)ö ·f FÒÒ" canadian balsamj ÒÏ~ ' bò¢ ò î .. 10 cm2 ×1 cm×10 10 cm2 ×10 cm 10 cm2 ×1cm×10 10 cm2 ×10cm. 3. Ö 5 V. 5% rose bengal formalin sol. 5% formalin 5% rose bengal formalin sol. 5% formalin 5% rose bengal formalin sol. 5% formalin 5% rose bengal formalin sol. 5% formalin 5% rose bengal formalin sol. 5% formalin 5% formalin 5% rose bengal formalin sol. 5% formalin 5% rose bengal formalin sol. 5% formalin 5% rose bengal formalin sol. 5% formalin 5% formalin 5% rose bengal formalin sol. 5% formalin 5% rose bengal formalin sol. 5% formalin. `÷`9 # º >` 2003j 7ú~ ÿã 131ê 30ª æFç~ §* 5êöB 17êræ~ 11B~ 7;&Bÿb Ò;6 'böB Â * 7;&Bÿb~ ÷ Wj Îv C 10B~ ÿb Â*~& (Table 2). Â* ª~ > º 2001jö "æöB >¯B ·>Ö¦(2002)~ 2001j Â* ª~ >f ?f 8j ¾æÚîb¾, 15B *ê~ ÿb Â*®~ 1996j¦V 2000jræ~ Ò Ö"ö j ² Ôf Â* ª~~ > (·> Ö¦ 1996, 1997, 1998, 1999, 2001).~ Î *êöB FÏ~(Nematodes), &BFÏ~(Sarcomastigophorans), &BW º'~(Harpacticoid copepods) 5 7'~~ F (Nauplius) Â*~&, Î~(Polychaetes), NÏ~ (Ostracods), jÿb~(Tardigrades) &*ê¢ 7 b Â*~&b, ÿ^ÿb~(Kinorhynchs)& §* 7ê öB, N~& §* 6êf 12êöB ¾æÒ, ·>.

(7) Min, W.-G. et al.. 258. Table 2. Number of meiobenthos at each stations of CCFZ in the Northeastern Pacific. (unit : No. of Inds./10 cm2) Station. N05 (%). N06 (%). N07 (%) 89 24. 60.5 54 16.3 32. 49.1 54 29.1 22. 58.7 58 55.2 41 23.9 27 25.7 27. 43.6 47 28.7 27. 5.4 5 3.4 13. 4.5 11 11.8 4. 12.0 4.3. 12.8 3 9.6 2. 3.7 2.5 2.5. Nematodes 41 Sarcomastigo- 20 phorans. 47.1 23.0. 64 50.8 35 27.8. Harpacticoids Nauplius. 16 2. 18.4 2.3. 11 6. 8.7 4.8. 8 5. Polychaetes Ostracods. 4 1. 4.6 1.1. 3 1. 2.4 0.8. 12 2. Tardigrades Kinorhynchs. 1. 1.1. 1. 0.8. 1. 0.8. Bivalves Loriciferans others Total. 3. 8.2 1.4. N08 (%). N09 (%). N10 (%) N12 (%) N13 (%) N14 (%) N16 (%) N17 (%). 9 7. 8.6 12 6.7 9. 2. 1.8. 1 1. 1.0 1.0. 2 1. 2.1 1.1 2. 2. 1.8. 1. 1.0. 1. 1.1. 1. 1.1. 1. 1.0. 58.0 4 33.3 2. 66.7 2 33.3 2 2 2. 25.0 17 37.8 25.0 18 40.0 25.0 5 25.0 5. 11.1 11.1. 2.0. 2 2.3 4 3.2 4 2.7 2 1.8 1 1.1 87 100.0 126 100.0 147 100.0 110 100.0 92 100.0 105 100.0 94 100.0 81 100.0 6 100.0 8 100.0 45 100.0. Ö¦(2002)~ 2001j öÒöB §* 10ê ¦"~ ;6öB ¾rb B {B ª~ ÿ7ÿb(Loriciferans) §* 10ê~ N10;6öBò BÒ>ÚB, §* 10ê ¦" ~ BB æöB~ ª ßWj Ò~& . ÿ7ÿb f jv' " 1983jö R.M. Kristensenö ~~ B ÒB ÿb^ (Giere 1993). ;6ê Â* ª~>º ; 6 N06, N10, N12öB 8B &Ë ô ¾æÒ, N14ö Bº 2B~ ª~ <ö Â*~æ p~b, ;6ö Bº 4~7B ;ê~ ª~ Â*~& (Table 2). &Ú &*êöB ôf ª~ Â*~º ·ç ¾æÒ . ' ;6ê ' ÷~ B&ê~ jNj , §* 17 ê¢ B Î *ê~ ;6öB FÏ~& Ö6~, 44~67%~ Ö6·çj ¾æî (Table 2). &Ë ¸f FÏ ~~ B jNj ;6f ;6 N14 *Ú~ 67%¢ Næ~ ®îb, r ;6 N07~ 61%, ;6 N09~ 59%î . v ® Ö6 ª~f &BF Ï~ 16~40%~ Ö6·çj &b, &Ë ¸f B jNj ;6f N17 *Ú~ 40%¢ Næ~ ®î , r ;6 N13~ 33%, ;6 N08~ 29%& . *Ú Â* ª~ 7öB FÏ~f &BFÏ~ v Næ~º jN 91%~50%¢ ¾æîº, ·>Ö¦ (1998)~ 70~82%, ·>Ö¦(1999)~ 56~73% º ¸, f (2000)~ 57~97%, ·>Ö¦(2002)~ 73~ 96%f jÝ Ö6·çj ¾æî . rb Ö6 &BW º'~~ ãÖº ;6 N05öB 18% &Ë ¸f B jNj ¾æîb, ;6 N06~ 13%, ;6 N09~ 12% ¾æÒ . ~ ;6öBº 4~11%~ Ö6·ç j &b¾, ;6 N14öBº BÚ~ &BW º' ~ê Â*~æ p~ . rb 7'~~ F Ö6 ~º ·çj 2~12%~ Wj¢ ¾æî .. ¢>'b ~ 7;&Bÿbº FÏ~¾ &BF Ï~, &BW º'~& çú~² Â* rJ^ ® (Shirayama 1983), ·>Ö¦~ 1996j~2001j~ CCFZ ~ Ò Ö"ê ¢ ÝA~ ® º ©j ' r ~ ª~ Wjº «*~ ¢>' &~ ª~ Wjf jÝ Ö"¢ " > ® . ;6ê Â*~º Ö6 ª~~ Wj¢ jv, &BFÏ~f &BW º'~~ BÚ>º ;6ê N& ìº >~, &Ë Ö6~º FÏ~~ Â* BÚ >~ N& ;6ê Wj~ N¢ &^Jº "º ö b (Table 2). *ê j÷ ò~ ' Ò;6~ [ 'b 3 cmÚöB Â* 7;&Bÿb~ *Ú BÚ>¢ , Ò;6 N07öB 147BÚ/10 cm &8j ¾æÚ, rb N06öB 126BÚ/10 cm ¢, ;6 N08f N10öB ' ' 110BÚ/10 cm , 105BÚ/10 cm ~ B&ê¢ ¾æî. . ~ ;6öBº 100BÚ/10 cm ~~ B&ê ¢ &, &Ë Ôf B&ê¢ ¾æÞ ;6f N14f N16b, 6, 8BÚ/10 cm ¾æÒ (Table 2; Fig. 2). f (2000)~ 1998j ¶òö ~~, §* 5êf 6êöB 300BÚ/10 cm ç~ ¸f B&ê¢ ¾æî, · >Ö¦(1999)~ 1999j ÒÖ"öº §* 5êf 6ê ; 6j B~º 2003j~ BÚ~ B&êf jvB £ 50BÚ/10 cm ;ê ç²~º B&ê¢ ©j J~, 2003j ~ 7;&Bÿb B&êº * >'b Ô² ¾æÒ . ãËf 1998jö B þÒö ~ 'Ëb [ ~ ¢NÖï~ jêê æÿ(·>Ö¦ 2001)" jÝ ãËj ¾æÚº 2. 2. 2. 2. 2. 2. 2. 2.

(8) Meiobenthic Community of Deep-sea Bed in the Northeastern Pacific. 259. B&ê¢ ¾æî, &BFÏ~f îR&æ ;6ê N& ² ¾æ¾æ p~ . 7;&Bÿbö ³~º ª ~öB jv' V& ö ³~º Î~º ;6 N07öB 12BÚ/10 cm ~ jv' ¸f B&ê¢ & b, ÿ^ÿb ;6 N07öBò 3BÚ/10 cm ~ & ê Â*j & . ~ ²> Â* ª~ §* 12ê ~~ &*ê~ ;6 öB Â*~º ·ç j & . Â*ª~ >f BÚ ~ B&ê~ &6ö B , §* 6~7êf 10~12ê Ò~ æöB 7;& Bÿb~ B ç&'b ·^ ©b . ¢> 'b ¦öB~ bî / ® B' .& æ(> 6,000 m ~~)öBº 10~100BÚ/10 cm ~ Ôf B &ê¢ ¾æÚ(Giere 1993), _ «¶~ / Ö ¾ >º æ ¯, ê.'b b2·Ê~ &B ¢Ú¾º jê~ " ''b &Ò ~ VÓ¾ êJbj 'Ò > ®º vª ®º æ öBº B&ê& 1,000BÚ/10 cm ö ~, Ïßæ " &6~ 7_ ¦"öBê ³¦ '·ª~ / Ú^ 7;&Bÿb~ &ê& ¸² ¾æ¾º ~ Ö"(Lambshead et al. 2001)öB, ¦öB / >º «¶W FVb >æ(sediment trap _f &î~ FV ê²¾ C î² 8~ ªCö ~)& ~ 7;&Bÿ b~ ª¢ ²Ö~º &Ë 7º ºb rJ^ ®. 2. 2. Fig. 2. Abundance of dominant meiofaunal groups at each station.. 2. 2. Fig. 3. Variation of meiofaunal density at the site of 5oN~10oN in the last several years.. Ö" [~ Öï æÿ ç7'b &Bê ö 'Ëj "îj &ËWj Ò (Fig. 3). 2003j~ Ôf B&êº §* 5êöB 10êræ~ ' ;6~ ï B&êº 1.11Ü10 BÚ/m B, Bï· öB >Úê 3.7Ü10 ~1.3Ü10 BÚ/m (Shirayama 1984a), §ï·öB~ 2.0Ü10 BÚ/m (Snider et al. 1984)f jvê ² Ôf º*~ 8¢ > ® . ª~ê * ;6öB &Ë ¸f FÏ~~ B &ê 8j ¾æÞ f ;6 N07 89BÚ/10 cm , r ;6 N06~ 64BÚ/10 cm , ;6N10~ 58BÚ /10 cm b ¾æÒ (Table 2). º ' ;6ê 7;&B ÿb~ *Ú Â* ·ç" ?f *çb, ' ;6~ 7; &Bÿb *Ú B&ê¢ ²Ö~º ª~ FÏ~ªj r > ® . r Ö6 &BFÏ~~ B &ê FÏ~f jÝ ·çj &º, ;6 N06ö B 35BÚ/10 cm , ;6 N08öB 32BÚ/10 cm *êö j~ ç&'b ¸² ¾æÒb¾, FÏ~~ ; 6ê B&êNö j~ j N¢ ¾æÚÚB, ;6ê B·çö N& ìº ©b . &B W º'~~ ãÖº ;6 N05öB 16BÚ/10 cm & 4. 2. 4. 6. 6. 2. 2. 2. 2. 2. 2. 2. 2. (Giere 1993; Neira et al. 2001; Shimanaga and Shirayama 2000; Sommer and Pfannkuche 2000; Soltwedel 2000; Danovaro et al. 2002). CCF 6. æ æ~ §* ê ¦"öB Î'ê ~f §'ê >~~ ãêF ®ÚB · ~& ò¾ >Z~º æ >N£[f *~ *ê j" '² pf ræ ;W>Ú ®, Ï Ö²[ê ö j pf ræ ;W>Ú ® V r^ö 1N Öï ¸² ¾æÒ º 6öB(·> Ö¦ 2001, 2002, 2004), 7;&Bb~ _ / ö ~æ, ßû ¾æ¾º §* 5êöB 7ê Ò ö *~ ~ 'böB ;6 ö j j" ¸f B&ê¢ ¾æÞ ©b ê (Fig. 4). ÿ¢æ öB ÒB &[ 'bÚ~ FVbï, ATP, Chl.a ª

(9) 7;&Bÿb~ B&ê~ ª

(10) NZ" FÒ ãËj & . ß® 'bÚ~ ;rÒj~ *ïj Ò~º ATP8(r=0.63)", R[b¦V ;B bW ê Jb·j æ~º Chl.a 8(r=0.64)f 7;&Bÿb~ B &êf F~>& 5%Ú~ ç&&ê& ¾æÒ (· >Ö¦ 2004). \8ê ª

(11) ß9 BÚ~ Vö V 7;&Bÿb~ ' ;6öB~ ª

(12) ·çj ÚÚ, Î ;6öB Ï 0.125 mm~ Ú.

(13) 260. Min, W.-G. et al.. Fig. 5. Number of meiobenthos depending on mesh sizes at each station.. Fig. 4. Abundance of Coccolithophores, Ciliates, Zooplankton, ATP (in sediment), Chl.a (in sediment), Meiobenthos at each station (KORDI 2004).. ¢ Û"~ 0.063 mmö æº V¢ &ê BÚ *Ú Vê B&êöB 41~51% &Ë ô ª

(14) ~ ®î (Fig. 5). rf Ï 0.063 mm~ Ú¢ Û"~ Ï 0.037 mm~ Ú Vö Îº V~ BÚ 11~36% ¸f jN ¾æ¾B jv' ·f V¢ & æº BÚ " ª

(15) ~º ©b . *êê Ú Ú, §* 5êöB 8êö º æFöB Ï 0.25 mm ç~ Úö Îº jv' V¢ &æº BÚ. ~ Wj& *ê~ Ã&ö V¢ 66 6²~º ãËj &, Ã&~ §* 13êöB &~ Wj¢ ¾æîb, §* 14êf 15öBº *& ¾æ¾æ p~. (Fig. 5). Vö V *êê ª

(16) ~ V& ¾æ ¾º Fº ~ ßWç BB _öB ö.æ >æ(Energy budget)¢ ' r Þ~ V& ·f © FÒ~V r^ö(Shirayama 1983), R[>~ ¢NÖï Ôf *êöB &[b />º '·bî BNb Þ V~ ²;z& >'>, ¢ NÖï ¸jB '·bî~ / ö &*ê öBº jv' V& BÚ Â*~º ©b Ò òB . ' ª~ê ÚÚ FÏ~º Î ;6öB Ú V 0.125 mm~ Ú¢ Û"~ 0.063 mmö æº V¢ &æº BÚ ~ BN Ö ¸f ©j " º(Fig. 5), ·>Ö¦(2001)~ 2000j ÒÖ"ê ÿ ¢ ·ç ¾æÒ . $ v ® Ö6 ª~ &B FÏ~ö ®ÚBê * ;6 'b [öB 0.125 mm~ Ú¢ Û"~ 0.063 mmö æº V¢ &æº BÚ. ô B~ ®î . rb Ö6 ª~ &BW º'~º ~ FÏ~ 5 &BFÏ~fº ², ² V 0.25 mm~ Ú¢ Û"~ 0.125 mm ö æº V¢ &æº BÚ ô ¾æÒ . $ *>'b 'b~ R[öB, 'b~ ¦[ö j. ² Þ~ V& BÚ Â*~º, º _ Âö j k öB _ jv' ±f R[j F^~V r^¢ ê (Sommer and Pfannkuche 2000). * ;6öB 7;&Bÿb~ 'b òö

(17) B &;&BÿbV~ bf ~ ¾æ¾æ p~b(·>Ö¦ 2004), Vê Â*çj 250~32 µm~ Úö Òº 7;&Bÿb ~ 90%ç j Næ~ ®î ..

(18) Meiobenthic Community of Deep-sea Bed in the Northeastern Pacific. 261. Fig. 6. Vertical distribution of dominant meiofaunal groups at each station.. >ç ª

(19) ß9 2003j 7ú §* 5êöB 17êö º ' *êê ; 6~ 7;&Bÿb~ 'b Ú >çª

(20) ¢ ÚÚ, * >'b 'b~ p& Ã&ö V¢ B&ê& * >'b 6²~º ãËj ¾æîb¾, pê · æz ·ç ¾æÒ (Fig. 6). 7;&Bÿb~ >çª

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(45) Meiobenthic Community of Deep-sea Bed in the Northeastern Pacific. ^^ò. Bÿ9. 1997. §ÿï·

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